Conceptualization, Y.G. and Z.L.; methodology, Y.P. and S.D.; software, Y.L. and Y.P.; data curation, L.X.; writing—original draft preparation, Y.P.; writing—review and editing, Y.P.; supervision, J.H.; funding acquisition, Y.G. All authors have read and agreed to the published version of the manuscript. This work was supported by the National Natural Science Foundation of China (No. 32072155), the Project of Education Department in Hunan (No. 18A101), and the double first-class construction project of Hunan Agricultural University (No. SYL2019013). C. elegans strains and E. coli OP50 were provided by the CGC, which is funded by the NIH Office of Research Infrastructure Programs (P40 OD010440).