Conceptualization, Y.Z. and Z.X.; methodology, J.P. and Y.D.; software, Y.T.; J.C., Z.H., Y.D. and T.H.; formal analysis, Y.T. and Z.H.; investigation, Y.D.; resources, J.P.; data curation, M.D.; writing—original draft preparation, J.P.; writing—review and editing, Y.T.; visualization, J.D.; supervision, Z.X.; project administration, Z.X.; funding acquisition, Z.X. and M.D. Y.T. and J.P. are the co-first authors. All authors have read and agreed to the published version of the manuscript. This study was supported by the National Natural Science Foundation of China (U20A20118), Natural Science Foundation of Hunan Province (2022JJ50264), and Research Foundation of Education Bureau of Hunan Province (21B0711).