The current study was supported by National Natural Science Foundation of China (Grant no. 31700332); Forestry Science and Technology Project of Hunan Province, China (Grant no. XKL201731); Science and Technology Project of Hunan Province, China (Grant nos. 2016TP2007, 2016TP1014); Special Financial Grant from the China Postdoctoral Science Foundation (Grant no. 2017T100782); Postdoctoral Science Foundation Project of Shaanxi Province, China (Grant no. 2016BSHEDZZ117). The funding agency was not involved in the design of the study, collection, analysis, and interpretation of data and in writing the manuscript.