This study was financially supported by the National Natural Science Foundation of China ( 51979104 ), the National Key Research and Development Program of China ( 2021YFC1910403 ), Natural Science Foundation of Hunan Province, China ( 2020JJ5019 ), the Research Foundation of Education Department of Hunan Province , China ( 21B0715 ), and the Science and Technology Innovation Program of Hunan Province ( 2020RC5008 ).